Materia Medica
Amaranth
Amaranthus blitum
Purple amaranth is a leafy annual eaten as a cooked green across the tropics and warm temperate regions. Its leaves are rich in β-carotene, vitamin C and minerals, and herbal traditions use it as a cooling, nourishing food.
What Is Amaranth?
Purple amaranth is a fast-growing leafy annual of the amaranth family, eaten as a cooked green. Its leaves are rich in β-carotene, vitamin C and minerals 5Reference 5Nutrients, minerals, pigments, phytochemicals and radical-scavenging activity in Amaranthus blitum — analytical characterisation of 16 genotypesView study →, and herbal traditions treat the plant as a cooling, nourishing food.
Amaranthus blitum is native to South America and has been introduced across much of the world 21Reference 21Amaranthus blitum L. — native and introduced distribution — species databaseView study →. It has been grown in Europe as a minor leaf vegetable 19Reference 19Amaranthus blitum Linnaeus — species description, habitat and distribution — floraView study →.
The leaf’s best-documented value is nutritional. In Kenyan preschool children, a recipe of sun-dried amaranth and cowpea leaves raised blood β-carotene, vitamin A and haemoglobin 3Reference 3Sun-dried cowpeas and amaranth leaves recipe improves β-carotene and retinol levels in serum and hemoglobin concentration among preschool children — human intervention studyView study →. Other research on the plant is in the test tube and in animals.
Amaranthus dubius, red spinach or spleen amaranth, is a different species with its own page. Amaranthus lividus is an older name for A. blitum 19,
How Is Amaranth Used?
Amaranth is used mainly as a cooked leafy vegetable. It also appears in traditional preparations such as decoctions, poultices, juices and soups.
The young leaves and shoots are steamed, sautéed, or added to broths and stews. Traditional practice gives the plant as a nourishing food during recovery from illness, digestive irritation or weakness.
In folk medicine, fresh leaf preparations are applied to inflamed skin, minor wounds and swelling. A. blitum has also been used for intestinal disorders, roundworm and bleeding 14Reference 14In vitroEvaluating anthelmintic, anti-platelet, and anti-coagulant activities, and identifying the bioactive phytochemicals of Amaranthus blitum L. — in vitroView study →.
Traditional Uses
Western Herbal Medicine
In Western herbal traditions amaranth is a nutritive and soothing herb. It is used for digestive irritation, constipation with dryness, and general nutritional support.
Its mineral-rich leaves were eaten as a strengthening food during convalescence and times of poor diet.
Traditional Chinese Medicine
In Traditional Chinese Medicine, amaranth species are considered cooling and are used to clear heat, cool the blood and promote urination. They are given for inflammatory conditions, summer heat and digestive irritation.
Some regional traditions also use amaranth for skin eruptions and damp-heat conditions.
Ayurvedic Medicine
Ayurveda classes amaranth as cooling, light and nourishing. It is used to pacify excess Pitta and to support digestion without heating.
The leaves are eaten during inflammatory states, digestive sensitivity, and conditions of excess heat or irritation.
Indications
- Nutritional depletion and weakness during recovery [cooked leaf; Western herbal medicine]
- Vitamin A deficiency [cooked leaf; food use]
- Digestive irritation [leaf; Western, Chinese and Ayurvedic medicine]
- Mild constipation with dryness [leaf; Western herbal medicine]
- Heat conditions and summer heat [leaf; TCM and Ayurveda]
- Mild urinary irritation [leaf; TCM]
- Inflamed skin and minor wounds [fresh leaf poultice; folk use]
- Roundworm [leaf; folk use]
- Bleeding [leaf; folk use] Caution: In the test tube, A. blitum extract reduced platelet clumping and slowed blood clotting, the opposite of a styptic effect 14.
Botany
Purple amaranth is a hairless annual herb of the amaranth family (Amaranthaceae). Its stems are 10–60 cm long and grow upright or sprawl along the ground, often branching from the base and sometimes forming mats. They are light green or purple 19,
The leaves are oval, 1–6 cm long, on stalks as long as the blade or longer. The tip of the leaf is distinctly notched, with a small point in the notch 19,
The tiny green flowers grow in clusters in the leaf axils and in slender spikes at the stem tips. The fruit is a small, flattened capsule, 1.2–3 mm long, that does not split open. Each holds a single shiny black or dark reddish-brown seed about 1–1.8 mm across 19,
Distribution
Purple amaranth is native to South America, in Peru, Bolivia, Brazil, Paraguay, Uruguay, northern Argentina and northern Chile 21Reference 21Amaranthus blitum L. — native and introduced distribution — species databaseView study →. It has been introduced across Europe, Africa, Asia, North America and the Pacific 21Reference 21Amaranthus blitum L. — native and introduced distribution — species databaseView study →.
It grows in fields, waste places, roadsides and other disturbed ground 19,
In Europe it was grown as a minor leaf vegetable, but that use is declining, and in many temperate countries it persists mostly as an occasional weed of greenhouses and gardens 19Reference 19Amaranthus blitum Linnaeus — species description, habitat and distribution — floraView study →.
Growing Conditions
Cultivation snapshot for Amaranth
- Fast-growing annual, full sun to part shade.
- Thrives in disturbed soils, gardens, roadsides and cultivated fields; tolerant of heat and drought.
Grow it yourself
Farm Wiki is our cultivation-focused sister site, also run by Justin Cooke.
Pharmacology & Research
The human research on Amaranthus blitum concerns the leaf as a source of provitamin-A carotenoids and iron in African diets, in children and young women 1,
The cardiovascular and cholesterol trials of “amaranth” used oil from the seeds of grain amaranths, rich in squalene, not the leaf 6,
- Best-supported: the leaf as a provitamin-A vegetable — a feeding study raised serum β-carotene, retinol and haemoglobin in children 3Reference 3Sun-dried cowpeas and amaranth leaves recipe improves β-carotene and retinol levels in serum and hemoglobin concentration among preschool children — human intervention studyView study →, though adding more leaf did not increase the iron absorbed 1Reference 1RCTA Higher Proportion of Iron-Rich Leafy Vegetables in a Typical Burkinabe Maize Meal Does Not Increase the Amount of Iron Absorbed in Young Women — randomised stable-isotope studyView study →.
-
Emerging, worth watching: in-vitro antioxidant activity from betalain pigments and flavonoids 4,
5Reference 4In vitroAmaranthus lividus leaf extract radical scavenging and phytochemical profile — in vitroView study →Reference 5Nutrients, minerals, pigments, phytochemicals and radical-scavenging activity in Amaranthus blitum — analytical characterisation of 16 genotypesView study → ; kidney protection in a rat oxidative-injury model 10Reference 10AnimalWater extract of A. lividus protects rat kidney against CCl₄ injury — animal studyView study →. -
Mechanistically thin: the cardiovascular evidence is for grain seed oil, not leaf 6,
7, ; neuroprotective, anthelmintic and antiplatelet effects are single in-vitro studies 14,8, 9Reference 6RCTAmaranth oil application for coronary heart disease and hypertension — randomised placebo-controlled trialView study →Reference 7Clinical trialMiroshnichenko, L. A., Zoloedov, V. I., Volynkina, A. P., & Kulakova, S. N. (2008). [Influence with amaranth and sunflower oils used in dietary therapy of patients with diebetes mellitus 2 types on parameters of carbohydrate and lipid metabolism] — clinical trial. Voprosy Pitaniia (in Russian). https://pubmed.ncbi.nlm.nih.gov/19227865/View study →Reference 8Clinical trialGonor, K. V., Pogozheva, A. V., Derbeneva, S. A., et al. (2006). [The influence of a diet with including amaranth oil on antioxidant and immune status in patients with ischemic heart disease and hyperlipoproteidemia] — comparative clinical study. Voprosy Pitaniia (in Russian). https://pubmed.ncbi.nlm.nih.gov/17313043/View study →Reference 9Clinical trialGonor, K. V., Pogozheva, A. V., Kulakova, S. N., et al. (2006). [The influence of diet with including amaranth oil on lipid metabolism in patients with ischemic heart disease and hyperlipoproteidemia] — comparative clinical study. Voprosy Pitaniia (in Russian). https://pubmed.ncbi.nlm.nih.gov/16862949/View study → 16Reference 14In vitroEvaluating anthelmintic, anti-platelet, and anti-coagulant activities, and identifying the bioactive phytochemicals of Amaranthus blitum L. — in vitroView study →Reference 16In vitroMethanol extracts of A. lividus and A. tricolor protect SH-SY5Y neuroblastoma cells against H₂O₂ — in vitroView study → . - The caveat: almost all non-nutritional evidence is preclinical, and several studies used other Amaranthus species.
Much of the “amaranth” literature is about other species or another plant part. The heart and cholesterol trials used grain-amaranth seed oil, and the best liver studies used A. spinosus, so their results do not carry over to A. blitum leaf.
1. Nutritive Value — Provitamin a & Iron
The leaf is a rich source of β-carotene, vitamin C and minerals 5Reference 5Nutrients, minerals, pigments, phytochemicals and radical-scavenging activity in Amaranthus blitum — analytical characterisation of 16 genotypesView study →. In preschool children in Machakos, Kenya, a cooked recipe of sun-dried amaranth and cowpea leaves raised serum β-carotene and retinol, and raised haemoglobin by 4.6%. Drying and cooking kept over 60% of the β-carotene 3Reference 3Sun-dried cowpeas and amaranth leaves recipe improves β-carotene and retinol levels in serum and hemoglobin concentration among preschool children — human intervention studyView study →.
In Burkina Faso, improved leaf sauces of amaranth or Ceylon spinach, eaten with the cereal paste tô twice a day, were estimated to supply 80–86% of young children’s average iron requirement and 90–170% of their vitamin A requirement 2Reference 2Contribution of leafy vegetable sauces to dietary iron, zinc, vitamin A and energy requirements in children and their mothers in Burkina Faso — dietary intake modellingView study →.
But in an iron-absorption study in 18 young women, meals with a sauce richer in amaranth leaf did not deliver more absorbed iron than a traditional sauce. The authors attributed this to the leaves’ high phenolic content 1Reference 1RCTA Higher Proportion of Iron-Rich Leafy Vegetables in a Typical Burkinabe Maize Meal Does Not Increase the Amount of Iron Absorbed in Young Women — randomised stable-isotope studyView study →.
Gap: the leaf’s iron is poorly absorbed, so it is more reliable as a vitamin-A vegetable than as an iron source.
2. Antioxidant Activity
The red and purple leaf forms owe their colour to betalain pigments, betacyanin and betaxanthin. In 16 A. blitum genotypes, pigment and phytochemical content went with antioxidant activity 5Reference 5Nutrients, minerals, pigments, phytochemicals and radical-scavenging activity in Amaranthus blitum — analytical characterisation of 16 genotypesView study →.
An A. lividus (A. blitum) leaf extract scavenged 75.6% of DPPH and 85.2% of superoxide radicals. Its main compounds were phytol and β-sitosterol, with gallic acid, caffeic acid, quercetin, vanillin and kaempferol 4Reference 4In vitroAmaranthus lividus leaf extract radical scavenging and phytochemical profile — in vitroView study →.
Gap: all of it is in vitro.
3. Cardiovascular (Seed Oil, Not Leaf)
In a randomised, placebo-controlled trial of 125 patients with coronary heart disease and hypertension, amaranth seed oil (3–18 mL a day, each 3 mL providing 100 mg squalene) lowered blood cholesterol and blood pressure 6Reference 6RCTAmaranth oil application for coronary heart disease and hypertension — randomised placebo-controlled trialView study →.
In Russian studies of patients with coronary heart disease and high blood lipids, diets with 100–600 mg of squalene a day from amaranth oil were compared. The 600 mg dose gave the largest improvement in cholesterol and triglycerides 9Reference 9Clinical trialGonor, K. V., Pogozheva, A. V., Kulakova, S. N., et al. (2006). [The influence of diet with including amaranth oil on lipid metabolism in patients with ischemic heart disease and hyperlipoproteidemia] — comparative clinical study. Voprosy Pitaniia (in Russian). https://pubmed.ncbi.nlm.nih.gov/16862949/View study →. The same dose gave the most favourable immune changes, and 200–400 mg the strongest antioxidant effect 8Reference 8Clinical trialGonor, K. V., Pogozheva, A. V., Derbeneva, S. A., et al. (2006). [The influence of a diet with including amaranth oil on antioxidant and immune status in patients with ischemic heart disease and hyperlipoproteidemia] — comparative clinical study. Voprosy Pitaniia (in Russian). https://pubmed.ncbi.nlm.nih.gov/17313043/View study →. Amaranth and sunflower oils in the diet also improved carbohydrate and lipid measures in type 2 diabetes 7Reference 7Clinical trialMiroshnichenko, L. A., Zoloedov, V. I., Volynkina, A. P., & Kulakova, S. N. (2008). [Influence with amaranth and sunflower oils used in dietary therapy of patients with diebetes mellitus 2 types on parameters of carbohydrate and lipid metabolism] — clinical trial. Voprosy Pitaniia (in Russian). https://pubmed.ncbi.nlm.nih.gov/19227865/View study →.
Squalene is a constituent of grain-amaranth seed oil, so these results cannot be claimed for A. blitum leaf.
Gap: no cardiovascular evidence exists for the leaf itself.
4. Organ Protection — Liver & Kidney
A water extract of A. lividus (a synonym of A. blitum), given to rats at 250 and 500 mg/kg for nine days, prevented the rise in kidney lipid peroxidation after carbon-tetrachloride injury and preserved catalase activity 10Reference 10AnimalWater extract of A. lividus protects rat kidney against CCl₄ injury — animal studyView study →.
The better-characterised liver-protection work used A. spinosus, a different species 11,
Gap: entirely preclinical and partly a different species, and CCl₄ injury is a toxicology model, not a human disease.
5. Other Preclinical Activity
A. blitum leaf extracts killed Trichinella spiralis larvae in the test tube at 1 mg/mL. The crude extract also reduced platelet aggregation (43% against 76% in control plasma) and slowed the extrinsic clotting pathway, and a newly identified coumaroyl-tryptophan compound was among the active constituents 14Reference 14In vitroEvaluating anthelmintic, anti-platelet, and anti-coagulant activities, and identifying the bioactive phytochemicals of Amaranthus blitum L. — in vitroView study →.
The larvicidal result matches the folk use against worms. The antiplatelet and anticoagulant results run the other way from the folk use for bleeding 14Reference 14In vitroEvaluating anthelmintic, anti-platelet, and anti-coagulant activities, and identifying the bioactive phytochemicals of Amaranthus blitum L. — in vitroView study →.
Methanol leaf extracts of A. lividus and A. tricolor protected human neuroblastoma cells against peroxide stress 16Reference 16In vitroMethanol extracts of A. lividus and A. tricolor protect SH-SY5Y neuroblastoma cells against H₂O₂ — in vitroView study →. In-vitro antidiabetic work (α-glucosidase inhibition and glucose uptake) used other species, A. cruentus and A. hybridus 15Reference 15In vitroIn vitro anti-diabetic activity and molecular docking studies of Amaranthus cruentus, Amaranthus hybridus and isolated compounds — in vitroView study →.
Mechanisms
The leaf’s antioxidant capacity comes from its betalain pigments, flavonoids and phenolic acids, shown in the test tube 4,
The cholesterol-lowering effects belong to squalene in grain-amaranth seed oil, a different plant part.
| Mechanism | Drives | Key compounds |
|---|---|---|
| Free-radical scavenging | antioxidant | betacyanin, betaxanthin |
| Radical scavenging; antiplatelet, larvicidal | antioxidantantiplateletanthelmintic | quercetin, kaempferol, gallic acid, caffeic acid |
| Retinol precursor | nutritive (vitamin A) | β-carotene |
| Cholesterol-synthesis modulation | cardiovascular (seed oil only) | squalene, tocotrienols |
Clinical Trials
No registered clinical trials of A. blitum leaf were identified. The human evidence is three nutrition studies. The cardiovascular trials used grain-amaranth seed oil, a different plant part and usually a different species.
| Completed | Planned | Terminated | Preclinical |
|---|---|---|---|
| 3(nutrition) | 0 | 0 | Many(in-vitro / animal) |
Last checked: July 2026.
Phytochemistry
Amaranthus blitum leaf is valued for a dense mix of pigments, vitamins and minerals rather than for a single active compound.
Its distinctive constituents are the betalain pigments betacyanin (red-purple) and betaxanthin (yellow), which colour the red and purple leaf forms. Across 16 genotypes, betacyanin ranged from 10.5 to 53.6 µg and betaxanthin from 18.5 to 58.4 µg per 100 g of fresh leaf 5Reference 5Nutrients, minerals, pigments, phytochemicals and radical-scavenging activity in Amaranthus blitum — analytical characterisation of 16 genotypesView study →.
The leaves are a strong source of β-carotene (442–1,282 µg/g fresh weight) and ascorbic acid (vitamin C, 123–1,848 µg/g) 5Reference 5Nutrients, minerals, pigments, phytochemicals and radical-scavenging activity in Amaranthus blitum — analytical characterisation of 16 genotypesView study →. They also contain magnesium, calcium, potassium, iron and zinc 5Reference 5Nutrients, minerals, pigments, phytochemicals and radical-scavenging activity in Amaranthus blitum — analytical characterisation of 16 genotypesView study →.
A leaf-extract profile of A. lividus identified phytol and β-sitosterol as major compounds, with the flavonoids quercetin and kaempferol and the phenolic acids gallic acid and caffeic acid 4Reference 4In vitroAmaranthus lividus leaf extract radical scavenging and phytochemical profile — in vitroView study →.
Grain-type amaranths concentrate the triterpene squalene in their seed oil. It is a seed constituent, not a leaf one 6Reference 6RCTAmaranth oil application for coronary heart disease and hypertension — randomised placebo-controlled trialView study →.
Constituent Summary
Pigment figures are µg per 100 g fresh leaf and vitamin figures µg per g fresh leaf, across 16 A. blitum genotypes 5Reference 5Nutrients, minerals, pigments, phytochemicals and radical-scavenging activity in Amaranthus blitum — analytical characterisation of 16 genotypesView study →. Leaf values vary widely with genotype, maturity and growing conditions.
Betalain2 compounds2 with data
Carotenoid1 compound1 with data
Vitamin1 compound1 with data
Sterol1 compoundno data
Triterpene1 compound1 with data
† The pigment that marks the red- and purple-leaved forms.
Dosage
Amaranth is a food. The leaves are eaten freely as a cooked vegetable, or taken as a decoction or infusion (1–2 cups daily) in traditional practice.
The standardised doses in the literature are for grain seed oil, a different plant part.
Nutrition (per 100 g, cooked leaves)
Cooked (boiled, drained) leaves are low in calories and a good source of vitamin C, vitamin A, calcium, iron, magnesium and folate:
| Nutrient | Per 100 g | % Daily Value |
|---|---|---|
| Energy | 21 kcal | 1% |
| Protein | 2.1 g | 5% |
| Carbohydrate | 4.1 g | 1% |
| Fat | 0.2 g | 0% |
| Vitamin C | 41 mg | 46% |
| Vitamin A | 139 µg RAE | 15% |
| Folate | 57 µg | 14% |
| Calcium | 209 mg | 21% |
| Iron | 2.3 mg | 13% |
| Magnesium | 55 mg | 13% |
| Potassium | 641 mg | 19% |
| Manganese | 0.86 mg | 37% |
Values for cooked, boiled, drained amaranth leaves; amaranth grain is a separate food with a different profile 18Reference 18Amaranth leaves, cooked, boiled, drained, without salt — food composition data (FDC ID 169202)View study →. The iron shown is total, not absorbed: the leaf’s iron is poorly absorbed 1Reference 1RCTA Higher Proportion of Iron-Rich Leafy Vegetables in a Typical Burkinabe Maize Meal Does Not Increase the Amount of Iron Absorbed in Young Women — randomised stable-isotope studyView study →.
Safety & Pregnancy
Amaranth leaf is safe as a food. Its cautions come from what the leaf takes up and stores: oxalate, nitrate and heavy metals.
- Safe as a food. No toxicity signal at normal dietary amounts.
- High oxalate. Amaranth stores calcium-oxalate crystals — limit it if prone to calcium-oxalate kidney stones 17Reference 17Effects of exogenous calcium on cadmium accumulation in amaranth (Amaranthus tricolor) — plant-physiology studyView study →.
- Nitrate accumulation. Leafy amaranth and its cooking water are not suitable for very young infants (methaemoglobinaemia risk).
- Heavy metals. Amaranth can take up cadmium — soil source matters for regular eaters 17Reference 17Effects of exogenous calcium on cadmium accumulation in amaranth (Amaranthus tricolor) — plant-physiology studyView study →.
Full Safety Detail
Food use. Amaranth is generally considered safe as a food, with no toxicity signal at dietary amounts.
Oxalate. Like spinach and chard, edible amaranth is rich in calcium-oxalate crystals 17Reference 17Effects of exogenous calcium on cadmium accumulation in amaranth (Amaranthus tricolor) — plant-physiology studyView study →. This matters for people prone to calcium-oxalate kidney stones.
Nitrate. Fast-growing leafy amaranth readily takes up nitrate, especially in heavily fertilised soil. High dietary nitrate in leafy greens can cause methaemoglobinaemia in infants, so amaranth leaf and its cooking water are not suitable for very young infants.
Cadmium. Edible amaranth (A. tricolor) is a potential cadmium hyperaccumulator 17Reference 17Effects of exogenous calcium on cadmium accumulation in amaranth (Amaranthus tricolor) — plant-physiology studyView study →, so soil source matters for anyone eating the leaf regularly. Cooking reduces oxalate and nitrate but does not remove accumulated metals.
Interactions and allergy. Herb–drug interactions have not been studied for A. blitum. Allergic reactions are rare but possible.
Amaranth leaf is safe to eat as a cooked vegetable in pregnancy. It is eaten that way in many cultures, with no specific safety signal at dietary amounts. Concentrated or medicinal preparations have not been studied in pregnancy or breastfeeding.
The usual leaf cautions still apply: cook it to reduce oxalate and nitrate, and mind the soil source for heavy metals.
References
- Cercamondi, C. I., et al. (2014). A Higher Proportion of Iron-Rich Leafy Vegetables in a Typical Burkinabe Maize Meal Does Not Increase the Amount of Iron Absorbed in Young Women — randomised stable-isotope study. Journal of Nutrition. https://doi.org/10.3945/jn.114.194670
- Icard-Vernière, C., Olive, F., Picq, C., & Mouquet-Rivier, C. (2015). Contribution of leafy vegetable sauces to dietary iron, zinc, vitamin A and energy requirements in children and their mothers in Burkina Faso — dietary intake modelling. Plant Foods for Human Nutrition. https://doi.org/10.1007/s11130-014-0462-5
- Nawiri, M. P., Nyambaka, H., & Murungi, J. I. (2013). Sun-dried cowpeas and amaranth leaves recipe improves β-carotene and retinol levels in serum and hemoglobin concentration among preschool children — human intervention study. European Journal of Nutrition. https://doi.org/10.1007/s00394-012-0360-2
- Durhan, B., Yalçın, E., Çavuşoğlu, K., & Acar, A. (2022). Amaranthus lividus leaf extract radical scavenging and phytochemical profile — in vitro. Scientific Reports. https://doi.org/10.1038/s41598-022-08421-8
- Sarker, U., & Oba, S. (2020). Nutrients, minerals, pigments, phytochemicals and radical-scavenging activity in Amaranthus blitum — analytical characterisation of 16 genotypes. Scientific Reports. https://doi.org/10.1038/s41598-020-59848-w
- Martirosyan, D. M., Miroshnichenko, L. A., Kulakova, S. N., et al. (2007). Amaranth oil application for coronary heart disease and hypertension — randomised placebo-controlled trial. Lipids in Health and Disease. https://doi.org/10.1186/1476-511X-6-1
- Miroshnichenko, L. A., Zoloedov, V. I., Volynkina, A. P., & Kulakova, S. N. (2008). [Influence with amaranth and sunflower oils used in dietary therapy of patients with diebetes mellitus 2 types on parameters of carbohydrate and lipid metabolism] — clinical trial. Voprosy Pitaniia (in Russian). https://pubmed.ncbi.nlm.nih.gov/19227865/
- Gonor, K. V., Pogozheva, A. V., Derbeneva, S. A., et al. (2006). [The influence of a diet with including amaranth oil on antioxidant and immune status in patients with ischemic heart disease and hyperlipoproteidemia] — comparative clinical study. Voprosy Pitaniia (in Russian). https://pubmed.ncbi.nlm.nih.gov/17313043/
- Gonor, K. V., Pogozheva, A. V., Kulakova, S. N., et al. (2006). [The influence of diet with including amaranth oil on lipid metabolism in patients with ischemic heart disease and hyperlipoproteidemia] — comparative clinical study. Voprosy Pitaniia (in Russian). https://pubmed.ncbi.nlm.nih.gov/16862949/
- Yilmaz-Ozden, T., Can, A., Karatug, A., et al. (2016). Water extract of A. lividus protects rat kidney against CCl₄ injury — animal study. Toxicology and Industrial Health. https://pubmed.ncbi.nlm.nih.gov/25415872/
- Zeashan, H., et al. (2009). Hepatoprotective and antioxidant activity of Amaranthus spinosus against CCl4 induced toxicity — in vitro. Journal of Ethnopharmacology. https://doi.org/10.1016/j.jep.2009.05.010
- Zeashan, H., et al. (2008). Hepatoprotective activity of Amaranthus spinosus in experimental animals — animal study. Food and Chemical Toxicology. https://doi.org/10.1016/j.fct.2008.08.013
- Salawu, Akindahunsi (2007). Ethanolic extracts of tropical vegetables including amaranth reduce serum AST/ALT after CCl₄ damage — animal study. Journal of Medicinal Food. https://doi.org/10.1089/jmf.2006.212
- Abdel-Moez, Sayed, Khalifa, et al. (2024). Evaluating anthelmintic, anti-platelet, and anti-coagulant activities, and identifying the bioactive phytochemicals of Amaranthus blitum L. — in vitro. BMC Complementary Medicine and Therapies. https://doi.org/10.1186/s12906-024-04478-2
- Nkobole, Ebenezer, Prinsloo, et al. (2025). In vitro anti-diabetic activity and molecular docking studies of Amaranthus cruentus, Amaranthus hybridus and isolated compounds — in vitro. BMC Complementary Medicine and Therapies. https://doi.org/10.1186/s12906-025-05169-2
- Amornrit, Santiyanont, et al. (2016). Methanol extracts of A. lividus and A. tricolor protect SH-SY5Y neuroblastoma cells against H₂O₂ — in vitro. Genetics and Molecular Research. https://doi.org/10.4238/gmr.15027562
- Xie, H., Liao, Z., Li, J., et al. (2023). Effects of exogenous calcium on cadmium accumulation in amaranth (Amaranthus tricolor) — plant-physiology study. Chemosphere. https://doi.org/10.1016/j.chemosphere.2023.138435
- USDA FoodData Central. Amaranth leaves, cooked, boiled, drained, without salt — food composition data (FDC ID 169202). https://fdc.nal.usda.gov/fdc-app.html#/food-details/169202/nutrients
- Mosyakin, S. L., & Robertson, K. R. (2003). Amaranthus blitum Linnaeus — species description, habitat and distribution — flora. In Flora of North America Editorial Committee (eds.), Flora of North America North of Mexico, Vol. 4. http://www.efloras.org/florataxon.aspx?flora_id=1&taxon_id=242302535
- Bao, B., Borsch, T., & Clemants, S. E. (2003). Amaranthus blitum Linnaeus — species description and distribution — flora. In Wu, Z. Y., & Raven, P. H. (eds.), Flora of China, Vol. 5. http://www.efloras.org/florataxon.aspx?flora_id=2&taxon_id=242302535
- Govaerts, R. (2026). Amaranthus blitum L. — native and introduced distribution — species database. The World Checklist of Vascular Plants (WCVP), Royal Botanic Gardens, Kew, via GBIF. https://www.gbif.org/species/8911082